S. G. Acinas, F. Rodriguez-valera, and C. Pedrós-alió, Spatial and temporal variation in marine bacterioplankton diversity as shown by RFLP fingerprinting of PCR amplified 16S rDNA, FEMS Microbiology Ecology, vol.24, issue.1, pp.27-40, 1997.
DOI : 10.1111/j.1574-6941.1997.tb00420.x

S. G. Acinas, J. Antón, and F. Valera, Diversity of freeliving and attached bacteria in offshore Western Mediterranean waters as depicted by analysis of genes encoding 16S rRNA, Appl. Environ. Microbiol, vol.65, pp.514-522, 1999.

K. R. Arrigo and G. L. Van-dijken, Annual cycles of sea ice and phytoplankton in Cape Bathurst polynya, southeastern Beaufort Sea, Canadian Arctic, Geophysical Research Letters, vol.45, issue.8, pp.10-1029, 2004.
DOI : 10.14430/arctic1413

K. R. Arrigo and G. L. Van-dijken, Secular trends in Arctic Ocean net primary production, Journal of Geophysical Research, vol.19, issue.9, p.10, 1029.
DOI : 10.1175/JCLI3767.1

F. Azam and F. Malfatti, Microbial structuring of marine ecosystems, Nature Reviews Microbiology, vol.16, issue.10, pp.782-791, 2007.
DOI : 10.2216/i0031-8884-16-3-253.1

F. Azam, D. C. Smith, G. F. Steward, and A. Hagstrom, Bacteriaorganic matter coupling and its significance for oceanic carbon cycling Microb, Ecol, vol.28, pp.167-17910, 1994.
DOI : 10.1007/bf00166806

L. Berdjeb, J. Ghiglione, and S. Jacquet, Bottom-Up versus Top-Down Control of Hypo- and Epilimnion Free-Living Bacterial Community Structures in Two Neighboring Freshwater Lakes, Applied and Environmental Microbiology, vol.77, issue.11, pp.3591-3599, 2011.
DOI : 10.1128/AEM.02739-10

M. Blais, J. Tremblay, A. D. Jungblut, J. Gagnon, J. Martin et al., Nitrogen fixation and identification of potential diazotrophs in the Canadian Arctic, Global Biogeochemical Cycles, vol.35, issue.5, p.302210, 1029.
DOI : 10.1046/j.1462-2920.2003.00451.x

D. Boeuf, M. T. Cottrell, D. L. Kirchman, P. Lebaron, J. et al., Summer community structure of aerobic anoxygenic phototrophic bacteria in the western Arctic Ocean, FEMS Microbiology Ecology, vol.85, issue.3, 2013.
DOI : 10.1111/1574-6941.12130

URL : https://hal.archives-ouvertes.fr/hal-01140647

B. J. Campbell and D. L. Kirchman, Bacterial diversity, community structure and potential growth rates along an estuarine salinity gradient, The ISME Journal, vol.58, issue.1, pp.10-220, 2012.
DOI : 10.4319/lo.2004.49.5.1620

URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC3526181

J. G. Caporaso, K. Bittinger, F. D. Bushman, T. Z. Desantis, G. L. Andersen et al., PyNAST: a flexible tool for aligning sequences to a template alignment, Bioinformatics, vol.26, issue.2, pp.266-26710, 2010.
DOI : 10.1093/bioinformatics/btp636

URL : http://www.ncbi.nlm.nih.gov/pmc/articles/PMC2804299

R. W. Corell, C. , and C. , Arctic Climate Impact Assessment (ACIA), in: Encyclopedia of Earth, National Council for Science and the Environment, 2010.

B. G. Crespo, T. Pommier, B. Fernández-gómez, and C. Pedrós-alió, Taxonomic composition of the particle-attached and free-living bacterial assemblages in the Northwest Mediterranean Sea analyzed by pyrosequencing of the 16S rRNA, MicrobiologyOpen, vol.5, issue.4, pp.10-1111, 2013.
DOI : 10.1371/journal.pcbi.1000352

B. C. Crump, J. A. Baross, and C. A. Simenstad, Dominance of particle-attached bacteria in the Columbia River estuary, USA, Aquatic Microbial Ecology, vol.14, pp.7-18, 1998.
DOI : 10.3354/ame014007

B. C. Crump, E. V. Armbrust, and J. A. Baross, Phylogenetic analysis of particle-attached and free-living bacterial communities in the Columbia river, its estuary, and the adjacent coastal ocean, Appl. Environ. Microbiol, vol.65, pp.3192-3204, 1999.

T. Dittmar and G. Kattner, The biogeochemistry of the river and shelf ecosystem of the Arctic Ocean: a review, Marine Chemistry, vol.83, issue.3-4, pp.103-12010, 2003.
DOI : 10.1016/S0304-4203(03)00105-1

D. Doxaran, J. Ehn, S. Bélanger, A. Matsuoka, S. Hooker et al., Optical characterisation of suspended particles in the Mackenzie River plume (Canadian Arctic Ocean) and implications for ocean colour remote sensing, Biogeosciences, vol.9, issue.8, pp.3213-322910, 2012.
DOI : 10.5194/bg-9-3213-2012

R. C. Edgar, Search and clustering orders of magnitude faster than BLAST, Bioinformatics, vol.26, issue.19, pp.2460-2461, 2010.
DOI : 10.1093/bioinformatics/btq461

E. A. Eloe, C. N. Shulse, D. W. Fadrosh, S. J. Williamson, E. E. Allen et al., Compositional differences in particle-associated and free-living microbial assemblages from an extreme deep-ocean environment, Environmental Microbiology Reports, vol.63, issue.4, pp.449-458, 2011.
DOI : 10.1016/j.tim.2006.06.008

A. Forest, M. Sampei, R. Makabe, H. Sasaki, D. G. Barber et al., The annual cycle of particulate organic carbon export in Franklin Bay (Canadian Arctic): Environmental control and food web implications, J. Geophys. Res.-Oceans, vol.113, pp.350510-1029, 2008.

A. Forest, M. Babin, L. Stemmann, M. Picheral, M. Sampei et al., Ecosystem function and particle flux dynamics across the Mackenzie Shelf Arctic Ocean): an integrative analysis of spatial variability and biophysical forcings, Biogeosciences Discuss, vol.95194, pp.10883-1096010, 2012.

C. A. Fuchsman, J. B. Kirkpatrick, W. J. Brazelton, J. W. Murray, and J. T. Staley, Metabolic strategies of free-living and aggregate-associated bacterial communities inferred from biologic and chemical profiles in the Black Sea suboxic zone, FEMS Microbiology Ecology, vol.78, issue.3, pp.586-603, 2012.
DOI : 10.1111/j.1574-6941.2011.01189.x

P. E. Galand, C. Lovejoy, J. Pouliot, M. E. Garneau, and W. F. Vincent, Microbial community diversity and heterotrophic production in a coastal Arctic ecosystem: A stamukhi lake and its source waters, Limnology and Oceanography, vol.53, issue.2, pp.813-823, 2008.
DOI : 10.4319/lo.2008.53.2.0813

E. Ortega-retuerta, Prokaryotic community structure and heterotrophic production in a river-influenced coastal arctic ecosystem, Aquat. Microb. Ecol, vol.42, pp.27-4010, 2006.

M. E. Garneau, W. F. Vincent, R. Terrado, and C. Lovejoy, Importance of particle-associated bacterial heterotrophy in a coastal Arctic ecosystem, Journal of Marine Systems, vol.75, issue.1-2, pp.185-197, 2009.
DOI : 10.1016/j.jmarsys.2008.09.002

J. F. Ghiglione and A. E. Murray, Pronounced summer to winter differences and higher wintertime richness in coastal Antarctic marine bacterioplankton, Environmental Microbiology, vol.45, issue.3, pp.617-629, 2012.
DOI : 10.1016/S0967-0637(98)00015-6

J. F. Ghiglione, M. Larcher, and P. Lebaron, Spatial and temporal scales of variation in bacterioplankton community structure in the NW Mediterranean Sea, Aquatic Microbial Ecology, vol.40, pp.229-24010, 2005.
DOI : 10.3354/ame040229

URL : https://hal.archives-ouvertes.fr/hal-00091839

J. F. Ghiglione, G. Mevel, M. Pujo-pay, L. Mousseau, P. Lebaron et al., Diel and Seasonal Variations in Abundance, Activity, and Community Structure of Particle-Attached and Free-Living Bacteria in NW Mediterranean Sea, Microbial Ecology, vol.10, issue.41, pp.217-231, 2007.
DOI : 10.1139/o59-099

J. F. Ghiglione, C. Palacios, J. C. Marty, G. Mével, C. Labrune et al., Role of environmental factors for the vertical distribution (0?1000 m) of marine bacterial communities in the NW Mediterranean Sea, Biogeosciences, vol.55194, pp.1751-176410, 1751.
URL : https://hal.archives-ouvertes.fr/hal-00330300

J. F. Ghiglione, P. Conan, and M. Pujo-pay, Diversity of total and active free-living vs. particle-attached bacteria in the euphotic zone of the NW Mediterranean Sea, FEMS Microbiology Letters, vol.299, issue.1, pp.9-21, 2009.
DOI : 10.1111/j.1574-6968.2009.01694.x

J. Ghiglione, P. E. Galand, T. Pommier, C. Pedrós-alió, E. W. Maas et al., Pole-to-pole biogeography of surface and deep marine bacterial communities, Proc. Natl
DOI : 10.1128/AEM.01541-09

J. A. Gilbert, D. Field, P. Swift, L. Newbold, A. Oliver et al., The seasonal structure of microbial communities in the Western English Channel, Environmental Microbiology, vol.95, issue.12, pp.3132-3139, 2009.
DOI : 10.1111/j.1462-2920.2009.02017.x

M. P. Ginige, P. Hugenholtz, H. Daims, M. Wagner, J. Keller et al., Use of Stable-Isotope Probing, Full-Cycle rRNA Analysis, and Fluorescence In Situ Hybridization-Microautoradiography To Study a Methanol-Fed Denitrifying Microbial Community, Applied and Environmental Microbiology, vol.70, issue.1, pp.588-596, 2004.
DOI : 10.1128/AEM.70.1.588-596.2004

L. D. Guo and R. W. Macdonald, Source and transport of terrigenous organic matter in the upper Yukon River: Evidence from isotope (delta C-13, Delta C-14, and delta N-15) composition of dissolved, colloidal, and particulate phases, Glob. Biogeochem. Cy, vol.20, pp.201110-1029, 2006.

L. R. Hodges, N. Bano, J. T. Hollibaugh, Y. , and P. L. , Illustrating the importance of particulate organic matter to pelagic microbial abundance and community structure??an Arctic case study, Aquatic Microbial Ecology, vol.40, pp.217-22710, 2005.
DOI : 10.3354/ame040217

J. T. Hollibaugh, P. S. Wong, and M. C. Murrell, Similarity of particle-associated and free-living bacterial communities in northern San Francisco Bay, California, Aquatic Microbial Ecology, vol.21, pp.103-11410, 2000.
DOI : 10.3354/ame021103

K. Holmfeldt, C. Dziallas, J. Titelman, K. Pohlmann, H. P. Grossart et al., along environmental gradients in the brackish northern Baltic Sea, Environmental Microbiology, vol.21, issue.8, pp.2042-2054, 2009.
DOI : 10.1002/j.1538-7305.1948.tb01338.x

S. M. Huse, D. M. Welch, H. G. Morrison, and M. L. Sogin, Ironing out the wrinkles in the rare biosphere through improved OTU clustering, Environmental Microbiology, vol.30, issue.7, pp.1889-1898, 2010.
DOI : 10.1101/gr.8.3.175

S. Ishii, M. Yamamoto, M. Kikuchi, K. Oshima, M. Hattori et al., Microbial Populations Responsive to Denitrification-Inducing Conditions in Rice Paddy Soil, as Revealed by Comparative 16S rRNA Gene Analysis, Applied and Environmental Microbiology, vol.75, issue.22, pp.7070-70781001481, 1128.
DOI : 10.1128/AEM.01481-09

C. T. Kellogg and J. W. Deming, Comparison of free-living, suspended particle, and aggregate-associated Bacterial and Archaeal communities in the Laptev Sea, Aquatic Microbial Ecology, vol.57, pp.1-1810, 2009.
DOI : 10.3354/ame01317

C. T. Kellogg, S. Carpenter, A. Renfro, A. Sallon, C. Michel et al., Evidence for microbial attenuation of particle flux in the Amundsen Gulf and Beaufort Sea: elevated hydrolytic enzyme activity on sinking aggregates, Polar Biology, vol.7, issue.12, 2007.
DOI : 10.5194/bg-7-1007-2010

P. F. Kemp, S. Lee, and J. Laroche, Estimating the growth rate of slowly-growing marine bacteria from RNA content, Appl. Environ . Microbiol, vol.59, pp.2594-2601, 1993.

L. Kernegger, I. Zweimuller, and P. Peduzzi, Effects of suspended matter quality and virus abundance on microbial parameters: experimental evidence from a large European river, Aquatic Microbial Ecology, vol.57, pp.161-17310, 2009.
DOI : 10.3354/ame01341

A. Lapoussiere, C. Michel, M. Starr, M. Gosselin, and M. Poulin, Role of free-living and particle-attached bacteria in the recycling and export of organic material in the Hudson Bay system, Journal of Marine Systems, vol.88, issue.3, pp.434-445, 2011.
DOI : 10.1016/j.jmarsys.2010.12.003

D. Lavoie, K. L. Denman, and R. W. Macdonald, Effects of future climate change on primary productivity and export fluxes in the Beaufort Sea, Journal of Geophysical Research, vol.32, issue.19, pp.10-1029, 2010.
DOI : 10.1007/978-3-642-18912-8_5

L. Fouest, V. Babin, M. Tremblay, and J. , The fate of riverine nutrients on Arctic shelves, Biogeosciences Discussions, vol.9, issue.10, pp.13397-13437, 2012.
DOI : 10.5194/bgd-9-13397-2012

URL : https://hal.archives-ouvertes.fr/hal-01089953

R. W. Macdonald, S. M. Solomon, R. E. Cranston, H. E. Welch, M. B. Yunker et al., A sediment and organic carbon budget for the Canadian Beaufort Shelf, Marine Geology, vol.144, issue.4, pp.255-27310, 1998.
DOI : 10.1016/S0025-3227(97)00106-0

M. Martinez-garcia, D. M. Brazel, B. K. Swan, C. Arnosti, P. S. Chain et al., Capturing Single Cell Genomes of Active Polysaccharide Degraders: An Unexpected Contribution of Verrucomicrobia, PLoS ONE, vol.215, issue.4, p.35314, 2012.
DOI : 10.1371/journal.pone.0035314.s012

A. Matsuoka, A. Bricaud, R. Benner, J. Para, R. Sempéré et al., Tracing the transport of colored dissolved organic matter in water masses of the Southern Beaufort Sea: relationship with hydrographic characteristics, Biogeosciences, vol.95194, pp.925-94010, 2012.

G. Mével, M. Vernet, M. Goutx, and J. F. Ghiglione, Seasonal to hour variation scales in abundance and production of total and particle-attached bacteria in the open NW Mediterranean Sea (0???1000 m), Biogeosciences, vol.5, issue.6, pp.1573-158610, 1573.
DOI : 10.5194/bg-5-1573-2008

M. C. Murrell, J. T. Hollibaugh, M. W. Silver, and P. S. Wong, Bacterioplankton dynamics in northern San Francisco Bay: Role of particle association and seasonal freshwater flow, Limnology and Oceanography, vol.44, issue.2, pp.295-308, 1999.
DOI : 10.4319/lo.1999.44.2.0295

I. Obernosterer, R. Lami, M. Larcher, N. Batailler, P. Catala et al., Linkage Between Bacterial Carbon Processing and the Structure of the Active Bacterial Community at a Coastal Site in the NW Mediterranean Sea, Microbial Ecology, vol.30, issue.3, pp.428-43510, 2010.
DOI : 10.1016/S0168-6496(00)00102-1

E. Ortega-retuerta, W. Jeffrey, J. Ghiglione, and F. Joux, Evidence of heterotrophic prokaryotic activity limitation by nitrogen in the Western Arctic Ocean during summer, Polar Biology, vol.32, issue.19, pp.785-794, 2012.
DOI : 10.1029/2005GL024216

E. Ortega-retuerta, W. H. Jeffrey, M. Babin, S. Bélanger, R. Benner et al., Carbon fluxes in the Canadian Arctic: patterns and drivers of bacterial abundance, production and respiration on the Beaufort Sea margin, Biogeosciences, vol.95194, pp.3679-369210, 2012.
URL : https://hal.archives-ouvertes.fr/hal-01253997

D. K. Perovich, The changing Arctic sea ice cover, Oceanography, pp.162-173, 2011.

B. J. Peterson, R. M. Holmes, J. W. Mcclelland, C. J. Vorosmarty, R. B. Lammers et al., Increasing River Discharge to the Arctic Ocean, Science, vol.298, issue.5601, pp.2171-2173, 2002.
DOI : 10.1126/science.1077445

URL : http://www.pik-potsdam.de/%7Estefan/Publications/Nature/peterson2002.pdf

C. Quince, A. Lanzen, R. J. Davenport, and P. J. Turnbaugh, Removing Noise From Pyrosequenced Amplicons, BMC Bioinformatics, vol.12, issue.1, pp.10-1186, 2011.
DOI : 10.1186/1471-2105-12-38

URL : http://doi.org/10.1186/1471-2105-12-38

J. Rath, K. Y. Wu, G. J. Herndl, and E. F. Delong, High phylogenetic diversity in a marine-snow-associated bacterial assemblage, Aquatic Microbial Ecology, vol.14, pp.261-26910, 1998.
DOI : 10.3354/ame014261

L. Riemann and M. Middelboe, Stability of bacterial and viral community compositions in Danish coastal waters as depicted by DNA fingerprinting techniques, Aquatic Microbial Ecology, vol.27, pp.219-23210, 2002.
DOI : 10.3354/ame027219

J. Rontani, B. Charriere, A. Forest, S. Heussner, F. Vaultier et al., Intense photooxidative degradation of planktonic and bacterial lipids in sinking particles collected with sediment traps across the, Canadian Beaufort ShelfArctic Ocean) Biogeosciences, vol.95194, pp.4787-480210, 2012.

Y. Shen, C. G. Fichot, and R. Benner, Dissolved organic matter composition and bioavailability reflect ecosystem productivity in the Western Arctic Ocean, Biogeosciences, vol.9, issue.12, pp.4993-500510, 2012.
DOI : 10.5194/bg-9-4993-2012

D. C. Smith and F. Azam, A simple, economical method for measuring bacterial protein synthesis rates in seawater using 3H- leucine, Mar. Microb. Food Webs, vol.6, pp.107-114, 1992.

D. C. Smith, M. Simon, A. L. Alldredge, and F. Azam, Intense hydrolytic enzyme activity on marine aggregates and implications for rapid particle dissolution, Nature, vol.359, issue.6391, pp.139-142, 1992.
DOI : 10.1038/359139a0

M. L. Sogin, H. G. Morrison, J. A. Huber, M. Welch, D. Huse et al., Microbial diversity in the deep sea and the underexplored "rare biosphere", Proc. Natl. Acad. Sci. USA, pp.12115-12120, 2006.
DOI : 10.1093/bioinformatics/18.11.1546

Y. Sun, R. D. Wolcott, and S. E. Dowd, Tag-Encoded FLX Amplicon Pyrosequencing for the Elucidation of Microbial and Functional Gene Diversity in Any Environment, Methods Mol. Biol, vol.733, pp.129-141, 2011.
DOI : 10.1007/978-1-61779-089-8_9

J. E. Tremblay and J. Gagnon, The effects of irradiance and nutrient supply on the productivity of Arctic waters: a perspective on climate change, Influence of Climate Change on the Changing Arctic and SubArctic Conditions, pp.73-93, 2009.
DOI : 10.1007/978-1-4020-9460-6_7

M. Troussellier, H. Schafer, N. Batailler, L. Bernard, C. Courties et al., Bacterial activity and genetic richness along an estuarine gradient (Rhone River plume, France), Aquatic Microbial Ecology, vol.28, pp.13-2410, 2002.
DOI : 10.3354/ame028013

C. Vallieres, L. Retamal, P. Ramlal, C. L. Osburn, and W. F. Vincent, Bacterial production and microbial food web structure in a large arctic river and the coastal Arctic Ocean, Journal of Marine Systems, vol.74, issue.3-4, pp.756-773, 2008.
DOI : 10.1016/j.jmarsys.2007.12.002

Q. Wang, G. M. Garrity, J. M. Tiedje, C. , and J. R. , Naive Bayesian Classifier for Rapid Assignment of rRNA Sequences into the New Bacterial Taxonomy, Applied and Environmental Microbiology, vol.73, issue.16, pp.5261-52671000062, 1128.
DOI : 10.1128/AEM.00062-07

P. Wassmann, C. M. Duarte, S. Agusti, and M. K. Sejr, Footprints of climate change in the Arctic marine ecosystem, Global Change Biology, vol.114, issue.2, pp.1235-1249, 2011.
DOI : 10.1029/2008JC005000

T. J. Williams, E. Long, F. Evans, M. Z. Demaere, F. M. Lauro et al., A metaproteomic assessment of winter and summer bacterioplankton from Antarctic Peninsula coastal surface waters, The ISME Journal, vol.12, issue.10, pp.1883-1900
DOI : 10.1073/pnas.1018221108

S. B. Wozniak, D. Stramski, M. Stramska, R. A. Reynolds, V. M. Wright et al., Optical variability of seawater in relation to particle concentration, composition, and size distribution in the nearshore marine environment at Imperial Beach, California, Journal of Geophysical Research, vol.43, issue.2, p.802710, 1029.
DOI : 10.5670/oceanog.2004.49