S. G. Acinas, L. A. Marcelino, V. Klepac-ceraj, and M. Polz, Divergence and Redundancy of 16S rRNA Sequences in Genomes with Multiple rrn Operons, Journal of Bacteriology, vol.186, issue.9, pp.2629-2635, 2004.
DOI : 10.1128/JB.186.9.2629-2635.2004

R. M. Atlas and R. Bartha, Biodegradation of petroleum in seawater at low temperatures, Canadian Journal of Microbiology, vol.18, issue.12, pp.1851-1855, 1972.
DOI : 10.1139/m72-289

S. Bordenave, M. S. Goñ-i-urriza, P. Caumette, and R. Duran, Effects of Heavy Fuel Oil on the Bacterial Community Structure of a Pristine Microbial Mat, Applied and Environmental Microbiology, vol.73, issue.19, pp.6089-6097, 2007.
DOI : 10.1128/AEM.01352-07

J. F. Braddock, M. L. Ruth, P. H. Catterall, J. L. Walworth, and K. A. Mccarthy, Enhancement and Inhibition of Microbial Activity in Hydrocarbon-Contaminated Arctic Soils:?? Implications for Nutrient-Amended Bioremediation, Environmental Science & Technology, vol.31, issue.7, 1997.
DOI : 10.1021/es960904d

J. R. Bragg, R. C. Prince, E. J. Harner, and R. M. Atlas, Effectiveness of bioremediation for the Exxon Valdez oil spill, Nature, vol.368, issue.6470, pp.413-418, 1994.
DOI : 10.1038/368413a0

O. G. Brakstad and A. G. Lødeng, Microbial Diversity during Biodegradation of Crude Oil in Seawater from the North Sea, Microbial Ecology 49, 94Á103, 2004.
DOI : 10.1007/s00248-003-0225-6

O. G. Brakstad, I. Nonstad, L. Faksness, and P. J. Brandvik, Responses of Microbial Communities in Arctic Sea Ice After Contamination by Crude Petroleum Oil, Microbial Ecology, vol.41, issue.3, pp.540-552, 2008.
DOI : 10.1139/f78-102

J. Brosius, T. J. Dull, D. D. Sleeter, and H. F. Noller, Gene organization and primary structure of a ribosomal RNA operon from Escherichia coli, Journal of Molecular Biology, vol.148, issue.2, pp.107-127, 1981.
DOI : 10.1016/0022-2836(81)90508-8

S. Cappello, G. Caruso, D. Zampino, L. S. Monticelli, G. Maimone et al., Microbial community dynamics during assays of harbour oil spill bioremediation: a microscale simulation study, Journal of Applied Microbiology, vol.20, issue.1, pp.184-194, 2007.
DOI : 10.1080/02757540410001728007

K. R. Clarke and R. M. Warwich, Primer-E (5) computer program, 2001.

F. Coulon, B. A. Mckew, A. M. Osborn, T. J. Mcgenity, and K. N. Timmis, Effects of temperature and biostimulation on oil-degrading microbial communities in temperate estuarine waters, Environmental Microbiology, vol.42, issue.1, pp.177-186, 2007.
DOI : 10.1099/ijs.0.02424-0

C. Delbè-s, J. J. Godon, and R. Moletta, Bacterial and archaeal 16S rDNA and 16S rRNA dynamics during an acetate crisis in an anaerobic digestor ecosystem, FEMS Microbiology Ecology, vol.35, issue.1, 2001.
DOI : 10.1016/S0168-6496(00)00107-0

U. Deppe, H. H. Richnow, and W. M. Antranikian, Degradation of crude oil by an arctic microbial consortium, Extremophiles, vol.48, issue.1, pp.461-470, 2005.
DOI : 10.1007/978-3-662-04643-2_15

D. Delille, Seasonal Variation of Bacteria in Sea Ice Contaminated by Diesel Fuel and Dispersed Crude Oil, Microbial Ecology, vol.33, issue.2, pp.97-105
DOI : 10.1007/s002489900012

D. Delille, A. Bassères, A. Dessommes, and C. Rosiers, Influence of daylight on potential biodegradation of diesel and crude oil in Antarctic seawater, Marine Environmental Research, vol.45, issue.3, pp.259-268, 1998.
DOI : 10.1016/S0141-1136(97)00129-3

D. Delille, E. Pelletier, A. Rodríguez-blanco, and J. F. Ghiglione, Effects of nutrient and temperature on degradation of petroleum hydrocarbons in sub-Antarctic coastal seawater, Polar Biology, vol.95, issue.10, 2009.
DOI : 10.1080/15320389509383486

D. Delille and E. Perret, Influence of temperature on the growth potential of Southern polar marine bacteria, Microbial Ecology, vol.22, issue.2, pp.117-123, 1989.
DOI : 10.1111/j.1365-2672.1977.tb00770.x

E. F. Delong, G. S. Wickham, and N. R. Pace, Phylogenetic stains: ribosomal RNA-based probes for the identification of single cells, Science, vol.243, 1989.

H. W. Ducklow, K. M. Myers, M. Erickson, J. F. Ghiglione, and A. E. Murray, Response of a summertime Antarctic marine ??bacterial community to glucose and ammonium enrichment, Aquatic Microbial Ecology, vol.64, issue.3, pp.205-220, 2011.
DOI : 10.3354/ame01519

M. Eriksson, J. O. Ka, and W. W. Mohn, Effects of Low Temperature and Freeze-Thaw Cycles on Hydrocarbon Biodegradation in Arctic Tundra Soil, Applied and Environmental Microbiology, vol.67, issue.11, pp.5107-5112, 2001.
DOI : 10.1128/AEM.67.11.5107-5112.2001

A. Rodríguez-blanco, Structure of total versus active bacterial communities exposed to hydrocarbons Citation: Polar Research, 2013.

G. Gentile, L. Giuliano, G. D-'auria, F. Smedile, M. Azzaro et al., Study of bacterial communities in Antarctic coastal waters by a combination of 16S rRNA and 16S rDNA sequencing, Environmental Microbiology, vol.55, issue.8, pp.2150-2161, 2006.
DOI : 10.1099/ijs.0.02424-0

B. Gerdes, R. Brinkmeyer, G. Dieckmann, and E. Helmke, Influence of crude oil on changes of bacterial communities in Arctic sea-ice, FEMS Microbiology Ecology, vol.53, issue.1, pp.129-139, 2005.
DOI : 10.1016/j.femsec.2004.11.010

J. F. Ghiglione, P. Conan, and M. Pujo-pay, Diversity of total and active free-living vs. particle-attached bacteria in the euphotic zone of the NW Mediterranean Sea, FEMS Microbiology Letters, vol.299, issue.1, pp.9-21, 2009.
DOI : 10.1111/j.1574-6968.2009.01694.x

J. F. Ghiglione, M. Larcher, and P. Lebaron, Spatial and temporal scales of variation in bacterioplankton community structure in the NW Mediterranean Sea, Aquatic Microbial Ecology, vol.40, pp.229-240, 2005.
DOI : 10.3354/ame040229

URL : https://hal.archives-ouvertes.fr/hal-00091839

J. F. Ghiglione and A. Murray, Pronounced summer to winter differences and higher wintertime richness in coastal Antarctic marine bacterioplankton, Environmental Microbiology, vol.45, issue.3, pp.617-629, 2012.
DOI : 10.1016/S0967-0637(98)00015-6

M. Givskov, L. Eberl, S. Moller, L. K. Poulsen, and S. Molin, Responses to nutrient starvation in Pseudomonas putida KT2442: analysis of general cross-protection, cell shape, and macromolecular content., Journal of Bacteriology, vol.176, issue.1, pp.7-14, 1994.
DOI : 10.1128/jb.176.1.7-14.1994

I. M. Head, D. M. Jones, and W. F. Rö-ling, Marine microorganisms make a meal of oil, Nature Reviews Microbiology, vol.308, issue.3, pp.173-182, 2006.
DOI : 10.1042/bst0230341

K. Hebenbrock, P. M. Williams, and B. L. Karger, Single strand conformational polymorphism using capillary electrophoresis with two-dye laser-induced fluorescence detection, Electrophoresis, vol.5, issue.1, 1995.
DOI : 10.1002/elps.11501601236

H. Hong, A. Pruden, and K. F. Reardon, Comparison of CE-SSCP and DGGE for monitoring a complex microbial community remediating mine drainage, Journal of Microbiological Methods, vol.69, issue.1, pp.52-64, 2007.
DOI : 10.1016/j.mimet.2006.11.016

D. M. Karl, The grounding of the Bahia Paraiso: Microbial ecology of the 1989 Antarctic oil spill, Microbial Ecology, vol.24, issue.1, pp.77-89, 1992.
DOI : 10.1007/BF00171972

P. F. Kemp, S. Lee, and J. Laroche, Estimating the growth rate of slowly growing marine bacteria from RNA content, Applied and Environmental Microbiology, vol.59, 1993.

D. L. Kirchman, X. A. Moran, and H. Ducklow, Microbial growth in the polar oceans ??? role of temperature and potential impact of climate change, Nature Reviews Microbiology, vol.19, pp.451-459, 2009.
DOI : 10.1007/s002489900003

J. A. Klappenbach, J. M. Dunbar, and T. M. Schmidt, rRNA Operon Copy Number Reflects Ecological Strategies of Bacteria, Applied and Environmental Microbiology, vol.66, issue.4, 2000.
DOI : 10.1128/AEM.66.4.1328-1333.2000

A. Ladousse and B. Tramier, Results of 12 Years of Research in Spilled Oil Bioremediation: INIPOL Eap 22, Proceedings of the 1991 Oil Spill Conference (Prevention, 1991.
DOI : 10.7901/2169-3358-1991-1-577

R. Lami, J. F. Ghiglione, Y. Desdevises, N. J. West, and P. Lebaron, Annual patterns of presence and activity of marine bacteria monitored by 16S rDNAÁ16S rRNA fingerprints in the coastal NW Mediterranean Sea, Aquatic Microbial Ecology, vol.54, 2009.

J. G. Leahy and R. R. Colwell, Microbial degradation of hydrocarbons in the environment, Microbiological Reviews, vol.54, pp.305-315, 1990.

D. H. Lee, Y. G. Zo, and S. J. Kim, Nonradioactive method to study genetic profiles of natural bacterial communities by PCR-single-strand-conformation-polymorphism, Applied and Environmental Microbiology, vol.62, pp.3112-3120, 1996.

D. K. Mills, K. Fitzgerald, C. D. Litchfield, and P. M. Gillevet, A comparison of DNA profiling techniques for monitoring nutrient impact on microbial community composition during bioremediation of petroleum-contaminated soils, Journal of Microbiological Methods, vol.54, issue.1, pp.57-74, 2003.
DOI : 10.1016/S0167-7012(03)00007-1

M. M. Moeseneder, J. M. Arrieta, and G. J. Herndl, A comparison of DNA- and RNA-based clone libraries from the same marine bacterioplankton community, FEMS Microbiology Ecology, vol.51, issue.3, pp.341-352, 2005.
DOI : 10.1016/j.femsec.2004.09.012

D. B. Nedwell, Effect of low temperature on microbial growth: lowered affinity for substrates limits growth at low temperature, FEMS Microbiology Ecology, vol.30, issue.2, pp.101-111, 1999.
DOI : 10.1111/j.1574-6941.1999.tb00639.x

O. Pelz, M. Tesar, R. M. Wittich, E. R. Moore, K. N. Timmis et al., Towards elucidation of microbial community metabolic pathways: unravelling the network of carbon sharing in a pollutant-degrading bacterial consortium by immunocapture and isotopic ratio mass spectrometry, Environmental Microbiology, vol.40, issue.2, pp.167-174, 1999.
DOI : 10.1080/10256019808036364

L. Pomeroy and W. J. Wiebe, Temperature and substrates as interactive limiting factors for marine heterotrophic bacteria, Aquatic Microbial Ecology, vol.23, pp.187-204, 2001.
DOI : 10.3354/ame023187

L. K. Poulsen, G. Ballard, and D. A. Stahl, Use of rRNA fluorescence in situ hybridization for measuring the activity of single cells in young and established biofilms, Applied and Environmental Microbiology, vol.59, 1993.

A. Rodríguez-blanco, V. Antoine, E. Pelletier, D. Delille, and J. F. Ghiglione, Effects of temperature and fertilization on total vs. active bacterial communities exposed to crude and diesel oil pollution in NW Mediterranean Sea, Environmental Pollution, vol.158, issue.3, pp.663-673, 2010.
DOI : 10.1016/j.envpol.2009.10.026

A. Rodríguez-blanco, J. F. Ghiglione, P. Catala, E. O. Casamayor, and P. Lebaron, Spatial comparison of total vs. active bacterial populations by coupling genetic fingerprinting and clone library analyses in the NW Mediterranean Sea, FEMS Microbiology Ecology, vol.67, issue.1, pp.30-42, 2009.
DOI : 10.1111/j.1574-6941.2008.00591.x

H. Schafer, L. Bernard, C. Courties, P. Lebaron, P. Servais et al., Microbial community dynamics in Mediterranean nutrient-enriched seawater mesocosms: changes in the genetic diversity of bacterial populations, FEMS Microbiology Ecology, vol.34, issue.3, pp.243-253, 2001.
DOI : 10.1016/S0168-6496(00)00102-1

M. A. Tumeo and A. E. Wolk, Assessment of the presence of oil-degrading microbes at McMurdo Station, Antarctic Journal of the United States, vol.29, pp.375-377, 1994.