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Mounaix for valuable support on the time-of-flight measurement and discussions in its interpretation We also thank E. Andressen and H. Rigneault for fruitful discussions and support, A. Malvache and R. Cossart for providing the brain slices, T. Gacoin and L. Meyer for providing the nanoKTP particles, and K. Vynck for fundamental input on the theoretical challenges. Funding: This work was supported by the FEMTO Network and contracts ANR-10-INBS-04-01 (France BioImaging infrastructure network, pp.11-17 ,